foc
《中国植物志》英文修订版
Flora of China
  FOC Vol.7 (2008)              | Family List  Cleomaceae  PDF     |  eFloras.org Link Font:+ Big | - Small
   10. Cleomaceae  
白花菜科   bai hua cai ke
  
Authors:Authors: Mingli Zhang & Gordon C. Tucker

  

Herbs [rarely shrubs], sometimes woody at base, producing mustard oils. Stems erect, sparsely or profusely branched, glabrous or glandular pubescent. Stipules scalelike or absent, caducous (petiolar spines sometimes present). Leaves alternate, spirally arranged, palmately compound; petiole often pulvinate; leaflets [1 or]3-7[-11]; leaflet blades with pinnate venation. Inflorescences racemes or corymbs or flowers solitary and axillary; peduncle present; bract present [or absent] at base of pedicels. Pedicel present; bracteoles absent. Flowers bisexual but sometimes appearing unisexual due to incomplete development, actinomorphic or slightly zygomorphic, rotate, crateriform, campanulate, or urceolate, hypogynous. Sepals 4, distinct or basally connate, persistent. Petals 4, distinct, imbricate, attached directly to receptacle; intrastaminal nectary-disk or glands present or sometimes absent. Stamens 6(-32); filaments free or basally adnate to gynophore (or along basal 1/3-1/2 in Gynandropsis); anthers dehiscing by longitudinal slits; pollen shed in single grains, 2-nucleate, commonly 3-colporate. Pistil 1, 2-carpellate; ovary superior; ovules 1 to many per locule, 2-tegmic, anatropous, placentation parietal; style 1, straight, short, thick; stigma 1, capitate, unlobed. Fruit an elongate capsule, ± dehiscent by lateral valves along their entire length [indehiscent or dehiscent schizocarp], usually stipitate from elongation of gynophore (lacking in Arivela). Seeds 1-10(-40) per capsule, tan, yellowish brown, or brown, cochleate-reniform, papillose or tuberculate, arillate or not; endosperm scanty or none but a persistent perisperm sometimes present.

Seventeen genera and ca. 150 species: worldwide in tropical and temperate regions; five genera and five species (three introduced) in China.

The generic reclassification of Cleomaceae followed here is based on studies of New World taxa by Iltis and Cochrane (Novon 17: 447-451. 2007). They recognized a number of smaller, more clearly delimited genera, based on seed anatomy and morphology, chromosome numbers, biogeographic analyses, and published molecular data. The resultant dismantling of the paraphyletic genus Cleome as it occurs in the New World is applicable in other regions, including Asia, because of the worldwide distribution of all the genera of the Cleomaceae.

The generic reclassification of Cleomaceae followed here is based on studies of New World taxa by Iltis and Cochrane (Novon 17: 447-451. 2007). They recognized a number of smaller, more clearly delimited genera, based on seed anatomy and morphology, chromosome numbers, biogeographic analyses, and published molecular data. The resultant dismantling of the paraphyletic genus Cleome as it occurs in the New World is applicable in other regions, including Asia, because of the worldwide distribution of all the genera of the Cleomaceae.

  
1Bract at base of pedicels simple; petals 1.5-4.5 cm; filaments 3-8.5 cm; anthers 6-10 mm; seeds 1.9-3.5 mm(2)
+Bract at base of pedicels 3-foliate or sometimes apical-most simple; petals 0.7-1.4 cm; filaments 0.5-3 cm; anthers 1-3 mm; seeds 1-1.6 mm(3)
   
2(1)Androgynophore 5-10 mm; gynophore 2-6 cm; nodal spines absent.2  Cleoserrata    西洋白花菜属
+Androgynophore absent; gynophore 4.5-8 cm; nodal spines present (prickles often present on petiole and major leaf veins).3  Tarenaya    醉蝶花属
   
3(1)Androgynophore 3-7 mm (in fruiting specimens look for separate scars from filaments distal to sepals or sepal scars).5  Gynandropsis    羊角菜属
+Androgynophore absent(4)
   
4(3)Stamens 6; sepals connate for 1/4-1/2 their length; leaflets 3; gynophore 3-12 mm.1  Cleome    白花菜属
+Stamens 14-25; sepals distinct; leaflets 3 or 5; gynophore absent.4  Arivela    黄花草属
   
   Lower Taxon
  • Arivela  Rafinesque  黄花草属
  • Cleome  Linnaeus  白花菜属
  • Gynandropsis  Candolle  羊角菜属
  • Tarenaya  Rafinesque  醉蝶花属
  • Cleoserrata  Iltis  西洋白花菜属